:root {
  --primary: #3CE88A;
  --primary-50: #0B2A22;
  --primary-900: #BEF9D6;
  --secondary: #16B89C;
  --accent: #FF8C6B;
  --neutral-50: #0E2622;
  --neutral-200: #1C4038;
  --neutral-500: #6E9A8E;
  --neutral-900: #E4F4ED;
  --bg: #06201F;
  --surface: #0E2E2B;
  --text: #E4F4ED;
  --text-muted: #A6CFC2;
  --success: #3CE88A;
  --warning: #F2C14E;
  --danger: #FF8C6B;
  --max-width: 72rem;
  --container-px: clamp(1.25rem, 4vw, 3rem);
  --font-heading: 'Bricolage Grotesque', system-ui, -apple-system, 'Segoe UI', sans-serif;
  --font-body: 'Figtree', system-ui, -apple-system, 'Segoe UI', sans-serif;
  --font-display: 'Bricolage Grotesque', 'Arial Black', 'Helvetica Neue', sans-serif;
  --fs-h1: clamp(2.125rem, 4.4vw, 3.375rem);
  --fs-h2: clamp(1.625rem, 3vw, 2.375rem);
  --fs-h3: 1.4375rem;
  --fs-body: 1.0625rem;
  --fs-small: 0.8125rem;
  --lh-heading: 1.14;
  --lh-body: 1.7;
  --radius-button: 22px 18px 24px 16px;
  --radius-card: 20px 28px 18px 26px;
  --on-primary: #111111;
}
*, *::before, *::after { box-sizing: border-box; }
* { margin: 0; }
html { -webkit-text-size-adjust: 100%; scroll-behavior: smooth; }
body { font-family: var(--font-body); font-size: var(--fs-body); line-height: var(--lh-body); color: var(--text); background: var(--bg); -webkit-font-smoothing: antialiased; }
h1, h2, h3, h4 { font-family: var(--font-heading); line-height: var(--lh-heading); font-weight: 700; color: var(--text); text-wrap: balance; }
h1 { font-size: var(--fs-h1); letter-spacing: -0.02em; }
h2 { font-size: var(--fs-h2); margin-top: 2.5rem; margin-bottom: 0.75rem; }
h3 { font-size: var(--fs-h3); margin-top: 1.75rem; margin-bottom: 0.5rem; }
p { margin-bottom: 1rem; max-width: 68ch; }
a { color: var(--primary); }
ul, ol { margin: 0 0 1rem 1.25rem; max-width: 68ch; }
li { margin-bottom: 0.35rem; }
strong { font-weight: 700; }
code { font-family: ui-monospace, "SF Mono", Menlo, monospace; background: var(--neutral-50); border: 1px solid var(--neutral-200); border-radius: 4px; padding: 0.1em 0.35em; font-size: 0.9em; }
blockquote { border-left: 3px solid var(--primary); padding-left: 1rem; color: var(--text-muted); margin: 0 0 1rem; }
img { max-width: 100%; height: auto; display: block; }
sup.cite a { text-decoration: none; color: var(--accent); font-weight: 600; }
.container { width: 100%; max-width: var(--max-width); margin-inline: auto; padding-inline: var(--container-px); }
.skip-link { position: absolute; left: -999px; }
.skip-link:focus { left: 1rem; top: 1rem; background: var(--surface); padding: 0.5rem 1rem; z-index: 100; }
.site-header { border-bottom: 1px solid var(--neutral-200); background: var(--surface); position: sticky; top: 0; z-index: 50; }
.site-header .container { display: flex; align-items: center; justify-content: space-between; min-height: 4rem; gap: 1rem; }
.site-brand { font-family: var(--font-display); font-weight: 800; font-size: 1.2rem; color: var(--text); text-decoration: none; }
.site-nav ul { display: flex; gap: 1.25rem; list-style: none; margin: 0; }
.site-nav a { color: var(--text-muted); text-decoration: none; font-size: var(--fs-small); font-weight: 600; }
.site-nav a:hover { color: var(--primary); }
.nav-toggle { display: none; background: none; border: 1px solid var(--neutral-200); border-radius: 6px; padding: 0.4rem 0.6rem; font-size: 1.1rem; cursor: pointer; color: var(--text); }
main { display: block; }
.hero { padding: clamp(2.5rem, 6vw, 5rem) 0 clamp(1.5rem, 4vw, 3rem); }
.hero .container { max-width: min(var(--max-width), 56rem); }
.eyebrow { font-family: var(--font-display); text-transform: uppercase; letter-spacing: 0.12em; font-size: var(--fs-small); font-weight: 700; color: var(--accent); margin-bottom: 0.75rem; }
.hero h1 { max-width: 22ch; }
.subheadline { font-size: 1.2rem; color: var(--text-muted); margin-top: 1rem; max-width: 60ch; }
.cta-row { display: flex; flex-wrap: wrap; gap: 0.75rem; margin-top: 1.75rem; }
.btn { display: inline-block; padding: 0.875rem 1.75rem; border-radius: var(--radius-button); font-weight: 700; text-decoration: none; font-size: var(--fs-small); transition: transform .15s ease, opacity .15s ease; }
.btn:hover { transform: translateY(-1px); }
.btn-primary { background: var(--primary); color: var(--on-primary); }
.btn-secondary { background: transparent; color: var(--primary); border: 1.5px solid var(--primary); }
article > section, main > article { padding-block: 0.5rem; }
.content { padding-block: 1rem 3rem; }
.hero-figure { margin: 1.5rem 0; }
.hero-figure img { border-radius: var(--radius-card); }
.card { background: var(--surface); border: 1px solid rule (#1C4038) with a faint 1px inner bloom #7CF5D8 top-edge highlight (the membrane's lit upper rim); discrete research-finding blocks use the membrane card, most short content uses NO card (organic negative space carries the separation); border-radius: var(--radius-card); padding: 1.75rem; }
.references-list { padding-left: 1.5rem; }
.references-list li { margin-bottom: 0.75rem; font-size: var(--fs-small); color: var(--text-muted); }
.references-list li a { word-break: break-word; }
.site-footer { background: var(--primary-900); color: var(--neutral-50); margin-top: 4rem; padding-block: 2.5rem; }
.site-footer a { color: var(--neutral-50); text-decoration: none; opacity: 0.85; }
.site-footer a:hover { opacity: 1; text-decoration: underline; }
.footer-disclaimer { font-style: italic; opacity: 0.8; margin-bottom: 1.5rem; max-width: 60ch; }
.footer-cols { display: grid; grid-template-columns: repeat(auto-fit, minmax(140px, 1fr)); gap: 1.5rem; }
.footer-cols h2 { font-size: var(--fs-small); text-transform: uppercase; letter-spacing: 0.08em; margin: 0 0 0.6rem; opacity: 0.7; }
.footer-cols ul { list-style: none; margin: 0; }
.footer-cols li { margin-bottom: 0.4rem; font-size: var(--fs-small); }
.copyright { margin-top: 1.75rem; font-size: var(--fs-small); opacity: 0.6; }
@media (max-width: 720px) {
  .nav-toggle { display: block; }
  .site-nav { display: none; position: absolute; top: 4rem; left: 0; right: 0; background: var(--surface); border-bottom: 1px solid var(--neutral-200); padding: 1rem var(--container-px); }
  .site-nav.open { display: block; }
  .site-nav ul { flex-direction: column; gap: 0.75rem; }
}
@media (prefers-reduced-motion: reduce) { * { scroll-behavior: auto !important; transition: none !important; } }
